n. The stone of an arch (typically the uppermost stone), which, being the last put in, is regarded as keying or locking the whole structure together; the stone at the apex of an arch.n. Hence A supporting principle; the chief element in a system; that upon which the remainder rests or depends.n. In chromolithography, the first stone on which the picture is drawn or photographed, to serve as an outline guide in preparing the other stones for the colors, a copy of the keystone being made on each stone for printing a single color. See lithography.n. In a Scotch lead-smelting furnace, a block of cast-iron used to close up the space at each end of the forestone, and to fill up the space between the forestone and the back part of the furnace