n. In machinery, a movable piece, generally of a cylindrical form, so fitted as to fill the sectional area of a tube, such as the barrel of a pump or the cylinder of a steam-engine, and capable of being driven alternately in two directions by pressure on one or the other of its sides.n. In musical wind-instruments of the trumpet family, one of the forms of valve whereby a crook is temporarily added to the tube and the pitch of the tones altered.n. In organ-building, a thumb-knob which may be pushed in like a piston, whereby some change in registration is pneumatically effected; a pneumatic coupler or combination knob.n. The central retractile part of the acetabulum or sucker of a cephalopod, whose action in producing a vacuum resembles that of the piston of an air-pump.