The American Heritage® Dictionary of the English Language, 4th Edition
n. The material or substance on which an enzyme acts.
n. Biology A surface on which an organism grows or is attached.
n. An underlying layer; a substratum.
n. Linguistics An indigenous language that contributes features to the language of an invading people who impose their language on the indigenous population.
n. A surface on which an organism grows or to which it is attached.
n. An underlying layer; a substratum.
n. A language that is replaced in a population by another language and that influences the language imposed on its speakers.
n. A metal which is plated with another metal which has different physical properties.
n. A surface to which a substance adheres.
n. The substance lining the bottom edge of an enclosure.
v. To strew or lay under.
the GNU version of the Collaborative International Dictionary of English
n. A substratum.
adj. Having very slight furrows.
v. To strew or lay under anything.
The Century Dictionary and Cyclopedia
n. A substratum.
To strew or lay under anything.
WordNet 3.0 Copyright 2006 by Princeton University. All rights reserved.
n. any stratum or layer lying underneath another
n. a surface on which an organism grows or is attached
n. an indigenous language that contributes features to the language of an invading people who impose their language on the indigenous population
n. the substance that is acted upon by an enzyme or ferment
Word Usage
"In a breakthrough discovery, we found that the target substrate is covalently modified by multiple moieties of APF-1, a reversible modification that renders the protein substrate susceptible to degradation."